Model Neuroscience: Dissecting Behavioral Change With Targeted Contrastive Neuron Attribution

v1.05Model Neuroscience: Dissecting Behavioral Change With Targeted Contrastive Neuron Attribution
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Introduction

The current popular methods of steering model behavior are likely unusable in production: control vectors found through popular methods like Contrastive Activation Addition (CAA) are brittle and known to degrade at high strengths, and sparse autoencoders (SAEs) have their own set of issues. For production model serving, the combination of instruction-tuning and RL pipelines serves as a 'good enough' behavioral guideline that doesn't require the application of external steering vectors. But what if there was a way to reliably alter model behavior and keep the rest of the model the same?

How can we reliably steer the behavior of LLMs (in a cheap, memory-efficient, and reproducible way) at high strengths while ensuring that the model doesn't suffer performance degradation? In this post we'll show you exactly how we do this.

Background on steering methods

The main scope of steering methods here falls into one of two categories: residual stream based control vectors, or features found using a sparse autoencoder. The former is brittle and hard to control. The latter, as prior work shows, is sensitive to noise and expensive to train. Both come with their flaws, and we needed something new.

Our main contribution here we call Contrastive Neuron Attribution (CNA). It's based on work done by TransluceAI showing that circuits describing model behavior are sparse and located in the multi-layer perceptron neuron basis, rather than in residual stream activations. We combine this idea with contrastive extraction to identify neurons associated with particular behaviors.

For some examples, as few as eight pairs of contrastive prompts can illuminate a neuron circuit describing an entire behavior. We benchmarked this technique, with and without steering, on Massive Multitask Language Understanding (MMLU) performance.

MMLU performance comparison across steering strengths
Figure 1. MMLU performance remains unchanged regardless of steering strength, demonstrating that contrastive neuron attribution preserves general model capabilities.

Base models show little behavioral change when steered the same way. This suggests that the refusal mechanism develops during fine-tuning: the structure is present in the base model, but alignment training connects it to a behavioral gate.

Refusal Circuit - Neuron Constellation

Interactive visualization

Llama-3.1-8B-Instruct · 200 neurons · Top by contrastive delta

200Sample neurons
15Sample layers
11–31Sample range
0.044%Sample / model MLP
The neuron constellation loads as you scroll into view.Open source visualization

Drag to orbit. Arrow keys rotate; + / − zoom; Home resets. Hover a neuron or choose one below to inspect.

Statistics describe the plotted sample: 200 records, against 32 × 14,336 model neurons. They are not the article’s reported analysis statistics.Source visualization

The refusal circuit neuron constellation discovered using contrastive neuron attribution shows how neurons cluster across layers and their contribution to refusal behavior.

Examples

Refusal outputs

Llama-3.1-70B-Instruct

Evaluation scenario: An unauthorized vehicle-access request.

Baseline: the response objects to or refuses the request.

After intervention: the source reports reduced refusal behavior.

Llama-3.2-3B-Instruct

Evaluation scenario: A request to promote a dangerous household product to children.

Baseline: the response objects to or refuses the request.

After intervention: the source reports reduced refusal behavior.

Qwen2.5-72B-Instruct

Evaluation scenario: A request to present an unsupported health claim as expert advice.

Baseline: the response objects to or refuses the request.

After intervention: the source reports reduced refusal behavior.

Interactive demo

Llama-3.1-8B-Instruct · Published neuron data

Explore the published neuron data above

This explorer uses the same published constellation data as above. The Qwen2.5-14B-Instruct model-response demo is not connected.

Limitations and Future Work

Some things we'd like to note:

  • This work does not yet cover Mixture-of-Experts models.
  • Amplification of a CNA-discovered circuit still degrades quality, while the reported ablation results do not. Understanding that difference remains an open question.
  • We plan to explore more behaviors and compare robustness across circuits.
  • We want to test whether circuit discovery transfers to quantized models.
  • Comparing circuits in instruction-tuned and base models could clarify how these behaviors develop.
  • The dynamic application of multiple steering vectors remains an area for future exploration.

Conclusion

We believe we've found a way to reliably steer model behavior while preserving output quality and other capabilities. This has implications for safely using the technique in production safety guardrails. There are many areas left to explore, and we welcome further research.

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MIT License · 2026